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Persat, Henri , Gaël Denys, Elsa Froufe, Giulia Secci-Petretto, and Steven Weiss Weiss. 2019. “Does Loire Grayling Represent A Distinct Species ?”. Frontiers In Marine Science 6. doi:10.3389/conf.fmars.2019.07.00070. http://www.frontiersin.org/Community/AbstractDetails.aspx?ABS_DOI=10.3389%2fconf.fmars.2019.07.00070.
Persat, Henri , Steven J Weiss, Elsa Froufe, Giulia Secci-Petretto, and Gaël Denys. 2019. “A Third European Species Of Grayling (Actinopterygii, Salmonidae), Endemic To The Loire River Basin (France), Thymallus Ligericus N. Sp.”. Cybium 43 (3): 233-238.
Persat, Henri , Gaël Denys, Agnès Dettai, and Philippe Keith. 2015. “The Freshwater Fish Fauna History Between Rhine And Ebro: General Considerations And Comments From On-Going Developments In Fish Taxonomy.”. Frontiers In Marine Science 2 (2015).